Short title Macroevolution and ecology drive multiple facets of tree diversityMartha Paola Barajas Barbosa*1,2(paolabarajas@gmail.com)Tiffany M. Knight1,3,4 Renske Onstein5,1 Jonathan Chase1,2 1 German Centre for Integrative Biodiversity Research (iDiv) Halle-Jena-Leipzig, Leipzig, Germany2 Department of Computer Science, Martin Luther University Halle-Wittenberg, Germany3 Department for Community Ecology, Helmholtz Centre for Environmental Research–UFZ, Leipzig, Germany4 Institute of Biology, Martin Luther University Halle-Wittenberg, Halle (Saale), Germany5 Naturalis Biodiversity Center, Darwinweg 2, 2333CR Leiden, the Netherlands*Corresponding author