Author’s contributions
EQ and JMR designed the study; EQ, RV, PYLB, GL did the sampling, CC carried out the DNA extraction, quality controls and PCRs under ALB supervision; SB, JM, RC and PYLB built the reference database and barcodes. CC, EQ, HL, PYLB and BB performed the analyses and interpreted the data; BB, CC and EQ led the writing and all the authors contributed to the manuscript and gave final approval for publication.