Leaf trait Abbreviation Mean Min Max CV (%)
Leaf chlorophyll content (SPAD) LCC 64.91 43.28 86.33 16.62
Leaf area (cm2) LA 18.44 7.62 41.36 48.81
Leaf thickness (mm) LT 0.55 0.36 0.82 23.64
Leaf volume (cm3) LV 1.03 0.34 2.76 61.17
Leaf fresh mass (g) LFM 1.05 0.36 2.62 60.00
Leaf saturated mass (g) LSM 1.18 0.43 2.90 59.32
Leaf dry mass (g) LDM 0.30 0.10 0.75 63.33
Leaf dry mass content (%) LDMC 25.38 14.42 34.34 16.58
Leaf density (g cm‒3) LD 0.29 0.14 0.40 20.69
Water saturation deficit (%) WSD 14.00 5.14 23.40 42.00
Leaf mass per area (g m−2) LMA 159.39 89.26 242.71 22.84