H. novo-guineensis:
Two of the species had no provenance-differentiation in Tleaf or Tcrit, but for differing reasons. For H. novo-guineensis , strong provenance-differentiation was observed in leaf width, leaf inclination angle, and the thermal time constant, all of which were higher for the upland provenance than the lowland provenance. However, the net effect of variation in these traits - wider leaves would increase Tleaf, but steeper leaf angles would reduce Tleaf - led to a lack of provenance-differentiation in Tmod that was supported by Tleaf. This highlights the importance of considering the suite of leaf thermal traits in combination.