Study area
The plant material for these studies of Origanum vulgare ssp.gracile was collected in 2022 in Tajikistan (Fig. 3) and is
deposited in the herbarium of the Central Siberian Botanical Garden
(NSK).
Population 1. 39°16’35.5”N 67°53’48.9”E. Zeravshan ridge. The upper
reaches of the river Mouserif. Belt of thermophilic junipers. Tree-shrub
layer: Juniperus semiglobosa Regel with an admixture ofJuniperus seravschanica Kom. Herb layer: Elymus repens(L.) Gould, Pseudoroegneria setulifera (Nevski) Á.Löve, Poa
versicolor Besser, Artemisia dracunculus L., Origanum
vulgare ssp. gracile , Phlomis cashmeriana Royle ex
Benth., Centaurea virgata ssp. squarrosa (Boiss.) Gugler,Ferula kokanica Regel & Schmalh., Arctium pseudarctium(Bornm.) Duist.
Population 2. 38°54’27.8”N 68°46’07.2”E. Hissar Ridge, a suburb of the
village Khoja-Obigarm. The upper boundary of the deciduous forest belt,
the right bank of the river. Tree-shrub layer: Acer platanoidesssp. turkestanicum (Pax) P.C. de Jong, Juniperus
seravschanica , Rosa kokanica (Regel) Regel ex Juz. Herb layer:Prangos pabularia Lindl., Origanum vulgare ssp.gracile , Hypericum perforatum L., Hypericum scabrumL., Ferula kuhistanica Korovin, Ferula jaeschkeana Vatke,Ziziphora pamiroalaica Juz., Dactylis glomerata L.
Population 3. 38°47’55.1”N 68°21’44.9”E. Hissar Ridge, a suburb of the
village Hakimi. Left bank of the river Karatag, the lower part of the
slope. Belt of thermophilic junipers. Tree-shrub layer: Juniperus
seravschanica , Fraxinus raibocarpa Regel, Prunus
bucharica (Korsh.) Hand.-Mazz., Acer pentapomicum J.L. Stewart,Lonicera nummulariifolia Jaub. & Spach. Herb layer:Festuca rupicola Heuff., Prangos pabularia , Hordeum
bulbosum L., Phlomis bucharica Regel, Ferula jaeschkeana ,Achillea filipendulina Lam., Lomelosia songarica (Schrenk)
Soják, Origanum vulgare ssp. gracile .
Population 4. 39°17’00.1”N 70°45’04.4”E. Karateginsky ridge, above the
mouth of the river Devonos, left tributary of the river Yarkhych.
Deciduous forest belt with shibleak elements. Tree-shrub layer:Acer platanoides ssp. turkestanicum , Cotoneaster
insignis Pojark., Lonicera nummulariifolia , Rosa
kokanica, Prunus cerasifera Ehrh. Herb layer: Dactylis
glomerata , Hordeum bulbosum , Ferula kokanica ,Ferula jaeschkeana , Salvia sclarea L., Salvia
turcomanica Pobed., Mentha longifolia (L.) L., Origanum
vulgare ssp. gracile .
Population 5. 38°40’40.2”N 69°29’49.2”E. Karategin range, a suburb of
the village Elok, river valley Gulhara. Deciduous forest belt with
fragments of thermophilic junipers. Tree-shrub layer: Rosa
achburensis Chrshan. Herb layer: Dactylis glomerata ,Phleum paniculatum Huds., Rumex paulsenianus Rech.f.,Elymus repens , Nepeta podostachys Benth., Nepeta
gontscharovii Kudrjasch., Handelia trichophylla (Schrenk)
Heimerl.