Study area
The plant material for these studies of Origanum vulgare ssp.gracile was collected in 2022 in Tajikistan (Fig. 3) and is deposited in the herbarium of the Central Siberian Botanical Garden (NSK).
Population 1. 39°16’35.5”N 67°53’48.9”E. Zeravshan ridge. The upper reaches of the river Mouserif. Belt of thermophilic junipers. Tree-shrub layer: Juniperus semiglobosa Regel with an admixture ofJuniperus seravschanica Kom. Herb layer: Elymus repens(L.) Gould, Pseudoroegneria setulifera (Nevski) Á.Löve, Poa versicolor Besser, Artemisia dracunculus L., Origanum vulgare ssp. gracile , Phlomis cashmeriana Royle ex Benth., Centaurea virgata ssp. squarrosa (Boiss.) Gugler,Ferula kokanica Regel & Schmalh., Arctium pseudarctium(Bornm.) Duist.
Population 2. 38°54’27.8”N 68°46’07.2”E. Hissar Ridge, a suburb of the village Khoja-Obigarm. The upper boundary of the deciduous forest belt, the right bank of the river. Tree-shrub layer: Acer platanoidesssp. turkestanicum (Pax) P.C. de Jong, Juniperus seravschanica , Rosa kokanica (Regel) Regel ex Juz. Herb layer:Prangos pabularia Lindl., Origanum vulgare ssp.gracile , Hypericum perforatum L., Hypericum scabrumL., Ferula kuhistanica Korovin, Ferula jaeschkeana Vatke,Ziziphora pamiroalaica Juz., Dactylis glomerata L.
Population 3. 38°47’55.1”N 68°21’44.9”E. Hissar Ridge, a suburb of the village Hakimi. Left bank of the river Karatag, the lower part of the slope. Belt of thermophilic junipers. Tree-shrub layer: Juniperus seravschanica , Fraxinus raibocarpa Regel, Prunus bucharica (Korsh.) Hand.-Mazz., Acer pentapomicum J.L. Stewart,Lonicera nummulariifolia Jaub. & Spach. Herb layer:Festuca rupicola Heuff., Prangos pabularia , Hordeum bulbosum L., Phlomis bucharica Regel, Ferula jaeschkeana ,Achillea filipendulina Lam., Lomelosia songarica (Schrenk) Soják, Origanum vulgare ssp. gracile .
Population 4. 39°17’00.1”N 70°45’04.4”E. Karateginsky ridge, above the mouth of the river Devonos, left tributary of the river Yarkhych. Deciduous forest belt with shibleak elements. Tree-shrub layer:Acer platanoides ssp. turkestanicum , Cotoneaster insignis Pojark., Lonicera nummulariifolia , Rosa kokanica, Prunus cerasifera Ehrh. Herb layer: Dactylis glomerata , Hordeum bulbosum , Ferula kokanica ,Ferula jaeschkeana , Salvia sclarea L., Salvia turcomanica Pobed., Mentha longifolia (L.) L., Origanum vulgare ssp. gracile .
Population 5. 38°40’40.2”N 69°29’49.2”E. Karategin range, a suburb of the village Elok, river valley Gulhara. Deciduous forest belt with fragments of thermophilic junipers. Tree-shrub layer: Rosa achburensis Chrshan. Herb layer: Dactylis glomerata ,Phleum paniculatum Huds., Rumex paulsenianus Rech.f.,Elymus repens , Nepeta podostachys Benth., Nepeta gontscharovii Kudrjasch., Handelia trichophylla (Schrenk) Heimerl.