Author Contributions
FY, LG and QJ designed the study. YL, CT, YZ, ZL, GW, PR, YH, XH, XC, LN, HS, MX and GL collected the specimens and performed the experiments. HX, BF, XFC, ZW, JY, YPH, TG, YJH and FY analysed the data. YL and JD wrote the manuscript. All authors read and approved the final manuscript.