Atmospheric CO2 concentrations are expected to continue increasing due to ongoing fossil fuel emissions, leading to innumerable climate impacts. These impacts are largely associated with the enhanced greenhouse effect, but recent work highlights that plant CO2 physiological responses can also strongly influence the climate. This study explores the impacts of plant responses on hydrological cycling at 2x preindustrial CO2 concentrations by analyzing simulations that separate plant physiology from climate responses to CO2 using the Community Earth System Model (CESM) versions 1 and 2. We find that CESM2 leaf area growth increases canopy evaporation, which offsets transpiration declines from reduced stomatal conductance, and dampens changes in precipitation, evapotranspiration, and runoff compared to CESM1 and a configuration of CESM2 with fixed leaf area. CESM2 better captures present-day leaf area magnitudes compared to observations but potentially over-estimates leaf area-CO2 sensitivity, highlighting the ongoing need to reduce plant allocation-related biases.

Plant stomata mediate the fluxes of both carbon and water between the land and the atmosphere. The ratio between photosynthesis and stomatal conductance (gs), or intrinsic water-use efficiency (iWUE), can be directly inferred from leaf or tree-ring carbon isotope composition. In many Earth system models, iWUE is inversely proportional and controlled by a parameter (g1M) in the calculation of gs. Here we examine how iWUE perturbations, setting g1M to the 5th (low) and 95th (high) percentile for each plant type based on observations, influence photosynthesis using coupled Earth System model simulations. We find that while lower iWUE leads to reductions in photosynthesis nearly everywhere, higher iWUE had a photosynthetic response that is surprisingly regionally dependent. Higher iWUE increases photosynthesis in the Amazon and central North America, but decreases photosynthesis in boreal Canada under fixed atmospheric conditions. However, the photosynthetic response to higher iWUE in these regions unexpectedly reverses when the atmosphere dynamically responds due to spatially differing sensitivity to increases in temperature and vapor pressure deficit. iWUE also influences the photosynthetic response to atmospheric CO2, with higher and lower iWUE modifying the total global response to elevated 2x preindustrial CO2 by 6.4% and -9.6%, respectively. Our work demonstrates that assumptions about iWUE in Earth system models significantly affect photosynthesis and its response to climate. Further, the response of photosynthesis to iWUE depends on which components of the model are included, therefore studies of iWUE impacts on historical or future photosynthesis can not be generalized across model configurations.

The insular Caribbean is a region influenced by Atlantic Ocean climate variability. Effects of low-frequency atmospheric circulation patterns on the precipitation of the Caribbean have been well documented. However, individual modes of variability are usually only considered in isolation. Here we analyse the combined and individual effects of the North Atlantic Oscillation (NAO) and the Atlantic Meridional Mode (AMM) on insular Caribbean precipitation. This work focuses on the Early Rainfall Season (ERS, April-July), which explains much of the interannual variability in precipitation for this region, from 1960-2016. Correlation analysis compare monthly NAO and AMM indices from the National Oceanic and Atmospheric Administration (NOAA) against monthly Caribbean precipitation from the Climate Research Unit (CRU) year-by-year climate variables by country. Sea surface temperature (SST) and sea level pressure (SLP) composites using NOAA data were also created to analyse regional patterns. Analysis of the results show that the NAO and AMM presented a correlation of opposite signs and affected the Eastern Caribbean (from Dominican Republic to Grenada) during ERS, resulting in precipitation anomalies above/below ± 10%. The combined and individual effects of NAO and AMM indicate that Feb-Mar NAO and AMM are significant correlated to May-Jun Eastern Caribbean precipitation anomalies. More frequent and consistent regional effects on precipitation anomalies, and more regionally spread and persistent SLP and SST were registered when both NAO and AMM occurred together in the previous winter. These results could be helpful in seasonal forecasting, by indicating whether a wetter or drier ERS would be expected based on the previous season NAO and AMM activity.