LT, LA, SLA, LDMC, LC, LN, LP, LN:LP and LC:LN represent leaf thickness, leaf area, specific leaf area, leaf dry matter content, leaf carbon content, leaf nitrogen content, leaf phosphorus content, leaf nitrogen-phosphorus ratio, and leaf carbon-nitrogen ratio, respectively.K : Blomberg’s K values, and significant values at p< 0.005 are shown in bold. All the trait data were log-transformed. p -values for traits with significant phylogenetic signals are highlighted in bold.