Author Contributions
BL, AK, ES, AT, ML, AG, FK, and AM designed and contributed to the original concept of the studies. BL and AG collected most of the specimens and related data, BL and ML obtained mtDNA data and prepared DNA libraries for ddRAD, AK conducted morphologic, diet and stable isotope analyses, AT conducted stable isotope analysis; ES conducted the most of bioinformatics, and BL, AK, ES, AG, FK and AM finalized the manuscript. All authors participated in project design, and read and approved the final manuscript.