Acknowledgements
We would like to appreciate our colleagues of the Key Laboratory for Sustainable Utilization of Marine Fisheries Resources of Yellow Sea Fisheries Research Institute, for their assistance on sample collection and helpful comments on the manuscript. This work was supported by the National Key R & D Program of China (No. 2019YFD0900403), Modern Agro-industry Technology Research System (No. CARS-48), National Natural Science Foundation of China (No. 31902367) and Central Public-interest Scientific Institution Basal Research Fund of CAFS (No. 2020TD46).