Acknowledgements
We thank Karin Köhl and the Green Team of the Max Planck Institute of Molecular Plant Physiology for taking excellent care of plants. We thank Maximillian Birkl, Christian Jorzig, Erik Jäsert and Gulmairam Sultanova for their help in plant phenotyping and Hao Tong for his help in GWA analysis. Jing Yu is acknowledged for her contribution in selecting accessions and in optimizing the N conditions. This work was supported by DFG priority programme 1819 (LA 3735/1-1 for G.T.D, P.K.P. and R.A.E.L. and LA 1472/4-1 for Z.N.) and the Max Planck Society (Z.N. and R.A.E.L.). In addition ARF, SA and Z.N. were supported by the European Union’s Horizon 2020 research and innovation programme, project PlantaSYST (SGA-CSA No. 739582 under FPA No. 664620).