Author contributions
GTD did all data analysis, ran GWAS, identified and characterized T-DNA
lines for the candidate genes, planned and performed the adaptability
experiment, planned and conducted the qRT-PCR. All figures were made by
GTD. PP conducted the initial screening and quantifications and
extraction of metabolites. NV measured the early rosette diameter and
helped PP with the screening. SA measured metabolite levels. ARF
supervised metabolomics measurements. RAEL and ZN planned the
experiments, RAEL supervised the experiments and ZN supervised data
analysis. RAEL, GTD and ZN wrote the manuscript. All authors have read
the final version of the manuscript.