Author contributions
GTD did all data analysis, ran GWAS, identified and characterized T-DNA lines for the candidate genes, planned and performed the adaptability experiment, planned and conducted the qRT-PCR. All figures were made by GTD. PP conducted the initial screening and quantifications and extraction of metabolites. NV measured the early rosette diameter and helped PP with the screening. SA measured metabolite levels. ARF supervised metabolomics measurements. RAEL and ZN planned the experiments, RAEL supervised the experiments and ZN supervised data analysis. RAEL, GTD and ZN wrote the manuscript. All authors have read the final version of the manuscript.