ACKNOWLEDGEMENTS
This research was supported by the National Natural Science Foundation of China (No. 41776162), the China Postdoctoral Science Foundation funded project (No. 2018M642712), the Youth Talent Support Program of the Laboratory for Marine Ecology and Environmental Science, the Pilot National Laboratory for Marine Science and Technology (Qingdao; LMEES-YTSP-2018-01-09), the Taishan Scholars Program (Distinguished Taishan Scholars), Two-Hundred Talents Plan of Yantai (Y839081021), the Youth Innovation Promotion Association CAS (2016196, 2019209), the Key Research Program (KFJ-STS-ZDTP-023), and the International Partners Program (133137KYSB20180069) of the Chinese Academy of Sciences.