The model
Dispersal models have been used to describe the implications of dispersal on the fitness of species that persist within a single habitat (Cohen, 1966, 1967; Fretwell and Lucas, 1970; Bulmer, 1984; Cohen and Levin, 1991), and dispersal ratios have been fixed (Hastings, 1983). Although, in some of these models, multiple sub-habitats are considered through the dispersal into or out of the stated sub-habitat, the models are not equipped to investigate the impact of dispersing into and out of multiple sub-habitats at once.