1Hsiung, Shih-Yi, 2Looy, Cindy,3Mander, Luke, and 4 Papper, Prahlada,1,2 Department of Integrative Biology and Museum of Paleontology, University of California, 3060 Valley Life Sciences Building #3140, Berkeley, California 94720, USA.3 College of Life and Environmental Sciences, University of Exeter, Prince of Wales Road, Exeter, Devon, EX4 4PS, UK.4 Department of Integrative Biology, University of California, 3060 Valley Life Sciences Building #3140, Berkeley, California 94720, USA.