Acknowledgements
We thank the Altizer lab for the use of flight mills, C. Gowler and A. Mongue for assistance with flight trials, and N. Barton and an anonymous reviewer for constructive feedback on a previous version of this manuscript. This work was supported by Emory University, National Science Foundation grants IOS-1557724, DEB-1754431 and IOS-1922720 to J.C.d.R., and National Science Foundation grant IOS-1452648 and National Institutes of Health grant GM108626 to M.R.K.