Figure 1. Spatial and social assortment . a , Palmyra Atoll Marine Reserve (red diamond) in the Central Pacific Ocean.b , Space use measured as the 50% utilisation distribution (UD) of sharks assigned to their respective colonies which were defined using community detection of individual movement networks (colours reflect colony groups in c ). c , Social networks and the distribution of weighted assortativity coefficients (\(r_{d}^{w}\)) for 10,000 random networks (boxes) and observed networks (red circles) across four years of shark telemetry data. Each node in the network represents an individual shark, with clusters showing closely associated dyadic pairs. Networks were all significantly, positively assorted by colonies, represented as different coloured nodes. No assortment is illustrated by blue dashed line. (p < 0.05*, p< 0.01** and p < 0.001***).